In an effort to understand the ecological role of elephants in forest environments, biologist Anika Raman conducted research in tropical regions where elephant populations had either remained stable, been reduced, or been completely removed due to habitat loss or human conflict. Raman observed that in areas where elephants were absent, the forest understory—the layer of vegetation beneath the forest canopy—became denser over time, inhibiting the growth of new saplings and reducing plant diversity. By contrast, in areas with stable elephant populations, the understory remained relatively open, allowing sunlight to reach the forest floor and enabling a greater variety of plant species to thrive. Based on these observations, Raman hypothesized that elephants act as "ecosystem engineers," helping to maintain plant diversity by trampling or feeding on fast-growing shrubs that would otherwise dominate the understory.
Which finding, if true, would most directly undermine Raman's hypothesis?